yes, direct url is working again.
That was it! Applied the one line change on all three servers here and the 60 query test goes from 20/60 to 60/60 on each of them.
For reference, temperror was running at 48%, 34% and 18% of inbound mail over the last 24 hours on those three. I will report back once enough new mail has come in to see where it lands.
Thanks for digging into this so quickly.